被子植物中具有性染色的類群

1、Homomorphic sex chromosomes indicates missing dataFamily(中文)Species（中文）Viability of YY genotypeSex determinationActinidiaceae 獼猴桃科Actinidia deliciosa 美味獼猴桃Active Y systemActinidiaceae 獼猴桃科A. chinensis 中華獼猴桃YesActive Y systemAmaranthaceae 莧科Acnida speciesActive Y systemAsparagaceae 天門冬科Asparagus offi

2、cinalis 石刁柏YesActive Y systemAsteraceae 菊科Antennaria dioica 山荻NoCaricaceae 番木瓜科Carica papaya 番木瓜NoActive Y systemCaricaceae 番木瓜科Vasconcellea species=(Vasconcellea pubescens？V. stipulata and V. cundinamarcensisCaryophyllaceae 石竹科Silene otites 蠅子草屬（歐洲） Chenopodiaceae 藜科Spinacia oleracea 菠菜YesActive Y

3、systemCucurbitaceae 葫蘆科Bryonia multiflora 甜瓜屬Cucurbitaceae 葫蘆科Ecballium elaterium 噴瓜Active Y systemDioscoreaceae 薯蕷科Dioscorea tokoro 山萆薢（bi xie）Active Y systemEuphorbiaceae 大戟科Mercurialis annua 西洋山靛YesActive Y systemRanunculaceae 毛茛科Thalictrum species YesRosaceae 薔薇科Fragaria speciesVitaceae 葡萄科Vitis

4、 speciesYesActive Y systemLife Form雌雄二倍體染色體數(shù)目傳粉Climbing shrubs Dioecydiploids, tetraploids and hexaploids2n=6x=174Climbing shrubs Dioecydiploid2n = 58*, 116*, 174*.Bees herbDioecy2n=32,34herbaceous perennial Dioecydiploid2n = 20herbaceous perennial Dioecydiploid2n = 28large tree-like plantpolygamous

5、diploid 2n = 18evergreen pachycaul shrubs or small treesdioecious and trioeciousdiploid 2n = 18herbaceous perennial dioecious. diploid 2n=24annual plantDioecy2n=12annual plantDioecy2n=20annual plantdioecious and monoecious n=12insectsherbaceous perennial lianasdioecious and monoecious diploid2n=20an

6、nual plantDioecyPolyploidy 2n=16?wind-pollinatedherbaceous perennialDioecy2n=14,2n=28,2n=42herbaceous perennialDioecyPolyploidy diploid 2n= 14herbaceous perennial lianasnormally dioeciousdiploid2n=38,40ReferenceFerguson, A. R., I. E. W. OBrien, and G. J. Yan.1997. Ploidy in Actinidia. Acta Horticult

7、urae 444: 6771；Testolin, R., G. Cipriani, and G. Costa. 1995. Sex segregation ratio and gender expression in the genus Actinidia. Sexual Plant Reproduction 8: 129132.Testolin, R., G. Cipriani, and G. Costa. 1995. Sex segregation ratio and gender expression in the genus Actinidia. Sexual Plant Reprod

8、uction 8: 129132.Charlesworth,D.,and D. Guttman. 1999. The evolution of dioecy and plant sex chromosome systems. In C. C. Ainsworth ed., Sex etermination in plants, 2549. BIOS Scientific Publishers, Oxford.UK; Costea, M. & D. DeMason (2001). Stem morphology and anatomy in Amaranthus L. (Amaranth

9、aceae)- Taxonomic significance. Journal of the Torrey Botanical Society 128(3): 254-281.Loptien, H. 1979. Identification of the sex chromosome pair in asparagus(Asparagus officinalis L.). Zeitschrift fur Pflanzenzuchtung 82: 162173.Ammal, E.& Kaul, B.Cytomorphological studies in autotetraploid A

10、sparagus officinalis L.Proceedings: Plant Sciences:Volume 65, Number 1,1-9, 1967.Charlesworth,D.,and D. Guttman. 1999. The evolution of dioecy and plant sex chromosome systems. In C. C. Ainsworth ed., Sex etermination in plants, 2549. BIOS Scientific Publishers, Oxford.UK.;RJ Bayer, GL Stebbins.Syst

11、ematic Botany (1987), 12(2): pp. 305-319,Chromosome Numbers, Patterns of Distribution, and Apomixis in Antennaria (Asteraceae: Inuleae)Horovitz, S., and H. Jimenez. 1967. Cruzamientos interespecficos e intergenericos en caricaceas y sus implicaciones fitotecnicas. Agronoma Tropical 17: 323343.Urasak

12、i, N.et al, 2002. A male and hermaphrodite specific RAPD marker for papaya (Carica papaya L.).Theoretical and Applied Genetics,104: 281-285.Horovitz, S., and H. Jimenez. 1967. Cruzamientos interespecficos e intergenericos en caricaceas y sus implicaciones fitotecnicas. Agronoma Tropical 17: 323343.C

13、 Balbontin et. al, 2010. VpAAT1, a Gene Encoding an Alcohol Acyltransferase, Is Involved in Ester Biosynthesis during Ripening of Mountain Papaya Fruit.J. Agric. Food Chem., 2010, 58 (8), pp 51145121.Charlesworth,D.,and D. Guttman. 1999. The evolution of dioecy and plant sex chromosome systems. In C

14、. C. Ainsworth ed., Sex etermination in plants, 2549. BIOS Scientific Publishers, Oxford.UK.;http:/nlbif.eti.uva.nl/bis/flora.php?menuentry=soorten&id=1966Warmake, H. E., and A. F.Blacklee. 1939. Sex mechanism inpolyploids of Melandrium album.Science 89: 391392.;Rebecca A.et. al, 1993. Flower De

15、velopmentin Dioecious Spinacia oleracea(Chenopodiaceae), A. J. bot.80(3):283-291.Charlesworth,D.,and D. Guttman. 1999. The evolution of dioecy and plant sex chromosome systems. In C. C. Ainsworth ed., Sex etermination in plants, 2549. BIOS Scientific Publishers, Oxford.UK.R. K. OYAMA et. al.,2009. A

16、 sex-linked SCAR marker in Bryonia dioica (Cucurbitaceae), a dioecious species with XY sex-determination and homomorphic sex chromosomes. Journal of Evolutionary Biology,Volume 22, Issue 1, pages 214224.Charlesworth,D.,and D. Guttman. 1999. The evolution of dioecy and plant sex chromosome systems. I

17、n C. C. Ainsworth ed., Sex etermination in plants, 2549. BIOS Scientific Publishers, Oxford.UK.;Reuyen Dukas,1987. Foraging behavior of three bee species in a natural mimicry system: female flowers which mimic male flowers in Ecballium elaterium (Cucurbitaceae) Oecologia Volume 74, Number 2, 256-263

18、;Thomas W. Whitaker, 1933.Cytological and Phylogenetic Studies in the Cucurbitaceae.Botanical Gazette Vol. 94, No. 4 , pp. 780-790.Martin and Oritz, 1963. Chromosome numbers and behaviorsin some species of Dioscorea. Cytologia 28: 96101; R Terauchi, G Kahl, 1999.Mapping of the Dioscorea tokoro genom

19、e: AFLP markers linked to sex. Genome,vol. 42, no4, pp. 752-762.Charlesworth,D.,and D. Guttman. 1999. The evolution of dioecy and plant sex chromosome systems. In C. C. Ainsworth ed., Sex etermination in plants, 2549. BIOS Scientific Publishers, Oxford.UK;S Hamdi et. al. 1987. Master Regulatory Gene

20、s, Auxin Levels, and Sexual Organogeneses in the Dioecious Plant Mercurialis annua.Plant Physiol. (1987) 85, 393-399.Charlesworth,D.,and D. Guttman. 1999. The evolution of dioecy and plant sex chromosome systems. In C. C. Ainsworth ed., Sex etermination in plants, 2549. BIOS Scientific Publishers, O

21、xford.UK.;Vincenzo Picci,1969.Ricerche Embriologiche Sul Genere Thalictrum. II. Embriologia e Cariologia Di Thalictrum Lucidum L. e di Thalictrum Minus L. Subsp. Minus.Plant Biosystems, Volume 103, Issue 6 , pages 475-483.Kihara, 1930. KaryologischeStudien an Fragaria mitbesonderer Berucksichtigung

22、derGeschlechtschromosomen.Cytologia1: 345357.Westergaard, 1958. The mechanism of sex determination in dioecious flowering plants. Advances in Genetics 9: 217281.Ferguson, A. R., I. E. W. OBrien, and G. J. Yan.1997. Ploidy in Actinidia. Acta Horticulturae 444: 6771；Testolin, R., G. Cipriani, and G. C

23、osta. 1995. Sex segregation ratio and gender expression in the genus Actinidia. Sexual Plant Reproduction 8: 129132.Testolin, R., G. Cipriani, and G. Costa. 1995. Sex segregation ratio and gender expression in the genus Actinidia. Sexual Plant Reproduction 8: 129132.Charlesworth,D.,and D. Guttman. 1

24、999. The evolution of dioecy and plant sex chromosome systems. In C. C. Ainsworth ed., Sex etermination in plants, 2549. BIOS Scientific Publishers, Oxford.UK; Costea, M. & D. DeMason (2001). Stem morphology and anatomy in Amaranthus L. (Amaranthaceae)- Taxonomic significance. Journal of the Tor

25、rey Botanical Society 128(3): 254-281.Loptien, H. 1979. Identification of the sex chromosome pair in asparagus(Asparagus officinalis L.). Zeitschrift fur Pflanzenzuchtung 82: 162173.Ammal, E.& Kaul, B.Cytomorphological studies in autotetraploid Asparagus officinalis L.Proceedings: Plant Sciences

26、:Volume 65, Number 1,1-9, 1967.Charlesworth,D.,and D. Guttman. 1999. The evolution of dioecy and plant sex chromosome systems. In C. C. Ainsworth ed., Sex etermination in plants, 2549. BIOS Scientific Publishers, Oxford.UK.;RJ Bayer, GL Stebbins.Systematic Botany (1987), 12(2): pp. 305-319,Chromosom

27、e Numbers, Patterns of Distribution, and Apomixis in Antennaria (Asteraceae: Inuleae)Horovitz, S., and H. Jimenez. 1967. Cruzamientos interespecficos e intergenericos en caricaceas y sus implicaciones fitotecnicas. Agronoma Tropical 17: 323343.Urasaki, N.et al, 2002. A male and hermaphrodite specifi

28、c RAPD marker for papaya (Carica papaya L.).Theoretical and Applied Genetics,104: 281-285.Horovitz, S., and H. Jimenez. 1967. Cruzamientos interespecficos e intergenericos en caricaceas y sus implicaciones fitotecnicas. Agronoma Tropical 17: 323343.C Balbontin et. al, 2010. VpAAT1, a Gene Encoding a

29、n Alcohol Acyltransferase, Is Involved in Ester Biosynthesis during Ripening of Mountain Papaya Fruit.J. Agric. Food Chem., 2010, 58 (8), pp 51145121.Charlesworth,D.,and D. Guttman. 1999. The evolution of dioecy and plant sex chromosome systems. In C. C. Ainsworth ed., Sex etermination in plants, 25

30、49. BIOS Scientific Publishers, Oxford.UK.;http:/nlbif.eti.uva.nl/bis/flora.php?menuentry=soorten&id=1966Charlesworth,D.,and D. Guttman. 1999. The evolution of dioecy and plant sex chromosome systems. In C. C. Ainsworth ed., Sex etermination in plants, 2549. BIOS Scientific Publishers, Oxford.UK

31、.R. K. OYAMA et. al.,2009. A sex-linked SCAR marker in Bryonia dioica (Cucurbitaceae), a dioecious species with XY sex-determination and homomorphic sex chromosomes. Journal of Evolutionary Biology,Volume 22, Issue 1, pages 214224.Charlesworth,D.,and D. Guttman. 1999. The evolution of dioecy and pla

32、nt sex chromosome systems. In C. C. Ainsworth ed., Sex etermination in plants, 2549. BIOS Scientific Publishers, Oxford.UK.;Reuyen Dukas,1987. Foraging behavior of three bee species in a natural mimicry system: female flowers which mimic male flowers in Ecballium elaterium (Cucurbitaceae) Oecologia

33、Volume 74, Number 2, 256-263;Thomas W. Whitaker, 1933.Cytological and Phylogenetic Studies in the Cucurbitaceae.Botanical Gazette Vol. 94, No. 4 , pp. 780-790.Martin and Oritz, 1963. Chromosome numbers and behaviorsin some species of Dioscorea. Cytologia 28: 96101; R Terauchi, G Kahl, 1999.Mapping o

34、f the Dioscorea tokoro genome: AFLP markers linked to sex. Genome,vol. 42, no4, pp. 752-762.Charlesworth,D.,and D. Guttman. 1999. The evolution of dioecy and plant sex chromosome systems. In C. C. Ainsworth ed., Sex etermination in plants, 2549. BIOS Scientific Publishers, Oxford.UK;S Hamdi et. al.

35、1987. Master Regulatory Genes, Auxin Levels, and Sexual Organogeneses in the Dioecious Plant Mercurialis annua.Plant Physiol. (1987) 85, 393-399.Charlesworth,D.,and D. Guttman. 1999. The evolution of dioecy and plant sex chromosome systems. In C. C. Ainsworth ed., Sex etermination in plants, 2549. B

36、IOS Scientific Publishers, Oxford.UK.;Vincenzo Picci,1969.Ricerche Embriologiche Sul Genere Thalictrum. II. Embriologia e Cariologia Di Thalictrum Lucidum L. e di Thalictrum Minus L. Subsp. Minus.Plant Biosystems, Volume 103, Issue 6 , pages 475-483.Westergaard, 1958. The mechanism of sex determinat

37、ion in dioecious flowering plants. Advances in Genetics 9: 217281.Ferguson, A. R., I. E. W. OBrien, and G. J. Yan.1997. Ploidy in Actinidia. Acta Horticulturae 444: 6771；Testolin, R., G. Cipriani, and G. Costa. 1995. Sex segregation ratio and gender expression in the genus Actinidia. Sexual Plant Re

38、production 8: 129132.Charlesworth,D.,and D. Guttman. 1999. The evolution of dioecy and plant sex chromosome systems. In C. C. Ainsworth ed., Sex etermination in plants, 2549. BIOS Scientific Publishers, Oxford.UK; Costea, M. & D. DeMason (2001). Stem morphology and anatomy in Amaranthus L. (Amar

39、anthaceae)- Taxonomic significance. Journal of the Torrey Botanical Society 128(3): 254-281.Loptien, H. 1979. Identification of the sex chromosome pair in asparagus(Asparagus officinalis L.). Zeitschrift fur Pflanzenzuchtung 82: 162173.Ammal, E.& Kaul, B.Cytomorphological studies in autotetraplo

40、id Asparagus officinalis L.Proceedings: Plant Sciences:Volume 65, Number 1,1-9, 1967.Charlesworth,D.,and D. Guttman. 1999. The evolution of dioecy and plant sex chromosome systems. In C. C. Ainsworth ed., Sex etermination in plants, 2549. BIOS Scientific Publishers, Oxford.UK.;RJ Bayer, GL Stebbins.

41、Systematic Botany (1987), 12(2): pp. 305-319,Chromosome Numbers, Patterns of Distribution, and Apomixis in Antennaria (Asteraceae: Inuleae)Horovitz, S., and H. Jimenez. 1967. Cruzamientos interespecficos e intergenericos en caricaceas y sus implicaciones fitotecnicas. Agronoma Tropical 17: 323343.Ur

42、asaki, N.et al, 2002. A male and hermaphrodite specific RAPD marker for papaya (Carica papaya L.).Theoretical and Applied Genetics,104: 281-285.Horovitz, S., and H. Jimenez. 1967. Cruzamientos interespecficos e intergenericos en caricaceas y sus implicaciones fitotecnicas. Agronoma Tropical 17: 3233

43、43.C Balbontin et. al, 2010. VpAAT1, a Gene Encoding an Alcohol Acyltransferase, Is Involved in Ester Biosynthesis during Ripening of Mountain Papaya Fruit.J. Agric. Food Chem., 2010, 58 (8), pp 51145121.Charlesworth,D.,and D. Guttman. 1999. The evolution of dioecy and plant sex chromosome systems.

44、In C. C. Ainsworth ed., Sex etermination in plants, 2549. BIOS Scientific Publishers, Oxford.UK.;http:/nlbif.eti.uva.nl/bis/flora.php?menuentry=soorten&id=1966Charlesworth,D.,and D. Guttman. 1999. The evolution of dioecy and plant sex chromosome systems. In C. C. Ainsworth ed., Sex etermination

45、in plants, 2549. BIOS Scientific Publishers, Oxford.UK.R. K. OYAMA et. al.,2009. A sex-linked SCAR marker in Bryonia dioica (Cucurbitaceae), a dioecious species with XY sex-determination and homomorphic sex chromosomes. Journal of Evolutionary Biology,Volume 22, Issue 1, pages 214224.Charlesworth,D.

46、,and D. Guttman. 1999. The evolution of dioecy and plant sex chromosome systems. In C. C. Ainsworth ed., Sex etermination in plants, 2549. BIOS Scientific Publishers, Oxford.UK.;Reuyen Dukas,1987. Foraging behavior of three bee species in a natural mimicry system: female flowers which mimic male flo

47、wers in Ecballium elaterium (Cucurbitaceae) Oecologia Volume 74, Number 2, 256-263;Thomas W. Whitaker, 1933.Cytological and Phylogenetic Studies in the Cucurbitaceae.Botanical Gazette Vol. 94, No. 4 , pp. 780-790.Martin and Oritz, 1963. Chromosome numbers and behaviorsin some species of Dioscorea. C

48、ytologia 28: 96101; R Terauchi, G Kahl, 1999.Mapping of the Dioscorea tokoro genome: AFLP markers linked to sex. Genome,vol. 42, no4, pp. 752-762.Charlesworth,D.,and D. Guttman. 1999. The evolution of dioecy and plant sex chromosome systems. In C. C. Ainsworth ed., Sex etermination in plants, 2549.

49、BIOS Scientific Publishers, Oxford.UK;S Hamdi et. al. 1987. Master Regulatory Genes, Auxin Levels, and Sexual Organogeneses in the Dioecious Plant Mercurialis annua.Plant Physiol. (1987) 85, 393-399.Charlesworth,D.,and D. Guttman. 1999. The evolution of dioecy and plant sex chromosome systems. In C.

50、 C. Ainsworth ed., Sex etermination in plants, 2549. BIOS Scientific Publishers, Oxford.UK.;Vincenzo Picci,1969.Ricerche Embriologiche Sul Genere Thalictrum. II. Embriologia e Cariologia Di Thalictrum Lucidum L. e di Thalictrum Minus L. Subsp. Minus.Plant Biosystems, Volume 103, Issue 6 , pages 475-

51、483.Ferguson, A. R., I. E. W. OBrien, and G. J. Yan.1997. Ploidy in Actinidia. Acta Horticulturae 444: 6771；Testolin, R., G. Cipriani, and G. Costa. 1995. Sex segregation ratio and gender expression in the genus Actinidia. Sexual Plant Reproduction 8: 129132.Charlesworth,D.,and D. Guttman. 1999. The

52、 evolution of dioecy and plant sex chromosome systems. In C. C. Ainsworth ed., Sex etermination in plants, 2549. BIOS Scientific Publishers, Oxford.UK; Costea, M. & D. DeMason (2001). Stem morphology and anatomy in Amaranthus L. (Amaranthaceae)- Taxonomic significance. Journal of the Torrey Bota

53、nical Society 128(3): 254-281.Loptien, H. 1979. Identification of the sex chromosome pair in asparagus(Asparagus officinalis L.). Zeitschrift fur Pflanzenzuchtung 82: 162173.Ammal, E.& Kaul, B.Cytomorphological studies in autotetraploid Asparagus officinalis L.Proceedings: Plant Sciences:Volume

54、65, Number 1,1-9, 1967.Charlesworth,D.,and D. Guttman. 1999. The evolution of dioecy and plant sex chromosome systems. In C. C. Ainsworth ed., Sex etermination in plants, 2549. BIOS Scientific Publishers, Oxford.UK.;RJ Bayer, GL Stebbins.Systematic Botany (1987), 12(2): pp. 305-319,Chromosome Number

55、s, Patterns of Distribution, and Apomixis in Antennaria (Asteraceae: Inuleae)Horovitz, S., and H. Jimenez. 1967. Cruzamientos interespecficos e intergenericos en caricaceas y sus implicaciones fitotecnicas. Agronoma Tropical 17: 323343.Urasaki, N.et al, 2002. A male and hermaphrodite specific RAPD m

56、arker for papaya (Carica papaya L.).Theoretical and Applied Genetics,104: 281-285.Horovitz, S., and H. Jimenez. 1967. Cruzamientos interespecficos e intergenericos en caricaceas y sus implicaciones fitotecnicas. Agronoma Tropical 17: 323343.C Balbontin et. al, 2010. VpAAT1, a Gene Encoding an Alcoho

57、l Acyltransferase, Is Involved in Ester Biosynthesis during Ripening of Mountain Papaya Fruit.J. Agric. Food Chem., 2010, 58 (8), pp 51145121.Charlesworth,D.,and D. Guttman. 1999. The evolution of dioecy and plant sex chromosome systems. In C. C. Ainsworth ed., Sex etermination in plants, 2549. BIOS

58、 Scientific Publishers, Oxford.UK.;http:/nlbif.eti.uva.nl/bis/flora.php?menuentry=soorten&id=1966Charlesworth,D.,and D. Guttman. 1999. The evolution of dioecy and plant sex chromosome systems. In C. C. Ainsworth ed., Sex etermination in plants, 2549. BIOS Scientific Publishers, Oxford.UK.R. K. O

59、YAMA et. al.,2009. A sex-linked SCAR marker in Bryonia dioica (Cucurbitaceae), a dioecious species with XY sex-determination and homomorphic sex chromosomes. Journal of Evolutionary Biology,Volume 22, Issue 1, pages 214224.Charlesworth,D.,and D. Guttman. 1999. The evolution of dioecy and plant sex c

60、hromosome systems. In C. C. Ainsworth ed., Sex etermination in plants, 2549. BIOS Scientific Publishers, Oxford.UK.;Reuyen Dukas,1987. Foraging behavior of three bee species in a natural mimicry system: female flowers which mimic male flowers in Ecballium elaterium (Cucurbitaceae) Oecologia Volume 74, Number 2, 256-263;Thomas W. Whitaker, 1933.C