細(xì)菌分類體系

細(xì)菌分類體系：1酸桿菌門(Acidobacteria)1.1酸桿菌綱(Acidobacteria)1.2全噬菌綱(Holophagae)2

放線菌門(Actinobacteria)2.1放線菌綱(Actinobacteria)2.2酸微菌亞綱(Acidimicrobidae)2.3放線菌亞綱(Actinobacteridae)2.4紅色桿菌亞綱(Rubrobacteridae)2.5球形桿菌亞綱(Sphaerobacteridae)3產(chǎn)水菌門(Aquificae)3.1產(chǎn)水菌綱(Aquificae)4擬桿菌門(Bacteroidetes)4.1擬桿菌綱(Bacteroidetes)4.2黃桿菌綱(Flavobacteria)4.3鞘脂桿菌綱(Sphingobacteria)4.4綱未定5衣原體門(Chlamydiae)5.1衣原體綱(Chlamydiae)6綠菌門(Chlorobi)6.1綠菌綱(Chlorobia)7綠彎菌門(Chloroflexi)7.1厭氧繩菌綱(Anaerolineae)7.2暖繩菌綱(Caldilineae)7.3綠彎菌綱(Chloroflexi)8產(chǎn)金菌門(Chrysiogenetes)8.1產(chǎn)金菌綱(Chrysiogenetes)9藍(lán)藻門(Cyanobacteria)9.1藍(lán)藻綱(Cyanobacteria)10脫鐵桿菌門(Deferribacteres)10.1脫鐵桿菌綱(Deferribacteres)11異常球菌-棲熱菌門(Deinococcus-Thermus)11.1異常球菌綱(Deinococci)12網(wǎng)團(tuán)菌門(Dictyoglomi)12.1網(wǎng)團(tuán)菌綱(Dictyoglomi)13纖維桿菌門(Fibrobacteres)13.1纖維桿菌綱(Fibrobacteres)14厚壁菌門(Firmicutes)（低G+C革蘭氏陽性菌）14.1芽孢桿菌綱(Bacilli)14.2梭菌綱(Clostridia)14.3熱石桿菌綱(Thermolithobacteria)15梭桿菌門(Fusobacteria)15.1梭桿菌綱(Fusobacteria)16芽單胞菌門(Gemmatimonadetes)16.1芽單胞菌綱(Gemmatimonadetes)17黏膠球形菌門(Lentisphaerae)17.1黏膠球形菌綱(Lentisphaerae)18硝化螺旋菌門(Nitrospirae)18.1硝化螺旋菌綱(Nitrospira)19浮霉菌門(Planctomycetes)19.1浮霉菌綱(Planctomycetacia)20海綿桿菌門(Poribacteria)21變形菌門(Proteobacteria)21.1α-變形菌綱(Alphaproteobacteria)21.2β-變形菌綱(Betaproteobacteria)21.3δ-變形菌綱(Deltaproteobacteria)21.4ε-變形菌綱(Epsilonproteobacteria)21.5γ-變形菌綱(Gammaproteobacteria)22螺旋體門(Spirochaetes)22.1螺旋體綱(Spirochaetes)23柔膜菌門(Tenericutes)23.1柔膜菌綱(Mollicutes)24熱脫硫桿菌門(Thermodesulfobacteria)24.1熱脫硫桿菌綱(Thermodesulfobacteria)25熱微菌門(Thermomicrobia)25.1熱微菌綱(Thermomicrobia)26熱袍菌門(Thermotogae)26.1熱袍菌綱(Thermotogae)27疣微菌門(Verrucomicrobia)27.1豐祐菌綱(Opitutae)27.2疣微菌綱(Verrucomicrobiae)28門未定28.1纖線桿菌綱(Ktedonobacteria)1酸桿菌門(Acidobacteria)1.1酸桿菌綱(Acidobacteria)酸桿菌目(Acidobacteriales)酸桿菌科(Acidobacteriaceae)酸桿菌屬(Acidobacterium)(Edaphobacter)(Terriglobus)1.2全噬菌綱(Holophagae)石鱉桿菌目(Acanthopleuribacterales)石鱉桿菌科(Acanthopleuribacteraceae)石鱉桿菌屬(Acanthopleuribacter)全噬菌目(Holophagales)全噬菌科(Holophagaceae)地發(fā)菌屬(Geothrix)全噬菌屬(Holophaga)2

放線菌門(Actinobacteria)2.1放線菌綱(Actinobacteria)2.2酸微菌亞綱(Acidimicrobidae)

酸微菌目(Acidimicrobiales)酸微菌亞目(Acidimicrobineae)酸微菌科(Acidimicrobiaceae)酸微菌屬(Acidimicrobium)(Iamiaceae)(Iamia)2.3放線菌亞綱(Actinobacteridae)

放線菌目(Actinomycetales)放線菌亞目(Actinomycineae)放線菌科(Actinomycetaceae)放線棒菌屬(Actinobaculum)放線菌屬(Actinomyces)隱秘桿菌屬(Arcanobacterium)(Falcivibrio)動彎桿菌屬(Mobiluncus)(Varibaculum)(Actinopolysporineae)(Actinopolysporaceae)(Actinopolyspora)(Catenulisporineae)(Actinospicaceae)(Actinospica)(Catenulisporaceae)(Catenulispora)棒桿菌亞目(Corynebacterineae)棒桿菌科(Corynebacteriaceae)(Bacterionema)(Caseobacter)棒桿菌屬(Corynebacterium)(Turicella)迪茨氏菌科(Dietziaceae)迪茨氏菌屬(Dietzia)分枝桿菌科(Mycobacteriaceae)分枝桿菌屬(Mycobacterium)

（含結(jié)核桿菌）諾卡氏菌科(Nocardiaceae)戈登氏菌屬(Gordonia)(Micropolyspora)(Millisia)諾卡氏菌屬(Nocardia)紅球菌屬(Rhodococcus)斯科曼氏菌屬(Skermania)(Williamsia)(Smaragdicoccus)(Segniliparaceae)(Segniliparus)束村氏菌科(Tsukamurellaceae)束村氏菌屬(Tsukamurella)弗蘭克氏菌亞目(Frankineae)酸熱菌科(Acidothermaceae)酸熱菌屬(Acidothermus)弗蘭克氏菌科(Frankiaceae)弗蘭克氏菌屬(Frankia)地嗜皮菌科(Geodermatophilaceae)芽球菌屬(Blastococcus)地嗜皮菌屬(Geodermatophilus)(Modestobacter)(Kineosporiaceae)(Cryptosporangium)(Kineococcus)(Kineosporia)(Nakamurellaceae)(Humicoccus)(Nakamurella)(Quadrisphaera)孢魚菌科(Sporichthyaceae)孢魚菌屬(Sporichthya)糖霉菌亞目(Glycomycineae)糖霉菌科(Glycomycetaceae)糖霉菌屬(Glycomyces)(Stackerbrandtia)微球菌亞目(Micrococcineae)獲山氏菌科(Beutenbergiaceae)獲山氏菌屬(Beutenbergia)喬治菌屬(Georgenia)薩勒河菌屬(Salana)博戈里亞湖菌科(Bogoriellaceae)博戈里亞湖菌屬(Bogoriella)短桿菌科(Brevibacteriaceae)短桿菌屬(Brevibacterium)纖維素單胞菌科(Cellulomonadaceae)纖維素單胞菌屬(Cellulomonas)厄氏菌屬(Oerskovia)(Tropheryma)(Dermabacteraceae)(Brachybacterium)(Dermabacter)(Dermacoccaceae)(Demetria)(Dermacoccus)(Kytococcus)(Dermatophilaceae)(Dermatophilus)(Kineosphaera)(Intrasporangiaceae)(Arsenicicoccus)(Humihabitans)(Intrasporangium)(Janibacter)(Knoellia)(Kribbia)(Lapillicoccus)(Ornithinicoccus)(Ornithinimicrobium)(Oryzihumus)(Serinicoccus)(Terrabacter)(Terracoccus)(Tetrasphaera)(Jonesiaceae)(Jonesia)(Microbacteriaceae)(Agreia)(Agrococcus)(Agromyces)(Aureobacterium)(Clavibacter)(Cryobacterium)(Curtobacterium)(Frigoribacterium)(Frondicola)(Gulosibacter)(Labedella)(Leifsonia)(Leucobacter)(Microbacterium)(Microcella)(Mycetocola)(Okibacterium)(Plantibacter)(Pseudoclavibacter)(Rathayibacter)(Rhodoglobus)(Salinibacterium)(Subtercola)(Yonghaparkia)(Zimmermannella)微球菌科(Micrococcaceae)(Acaricomes)節(jié)桿菌屬(Arthrobacter)(Citricoccus)(Kocuria)微球菌屬(Micrococcus)(Nesterenkonia)(Renibacterium)羅氏菌屬(Rothia)口腔球菌屬(Stomatococcus)劉志恒菌屬(Zhihengliuella)(Promicromonosporaceae)(Cellulosimicrobium)(Isoptericola)(Myceligenerans)(Promicromonospora)(Xylanibacterium)(Xylanimonas)(Rarobacteraceae)(Rarobacter)(Sanguibacteraceae)血桿菌屬(Sanguibacter)(Yaniaceae)(Yania)科未定

(Actinotalea)(Demequina)(Phycicoccus)(Ruania)微單孢菌亞目(Micromonosporineae)微單孢菌科(Micromonosporaceae)(Actinocatenispora)(Actinoplanes)(Amorphosporangium)(Ampullariella)(Asanoa)(Catellatospora)(Catenuloplanes)(Couchiolanes)(Dactylosporangium)(Krasilnikovia)(Longispora)(Luedemannella)微單孢菌屬(Micromonospora)(Pilimelia)(Planopolyspora)(Polymorphospora)(Salinispora)(Spirilliplanes)(Verrucosispora)(Virgisporangium)丙酸桿菌亞目(Propionibacterineae)(Nocardioidaceae)(Actinopolymorpha)(Aeromicrobium)(Friedmanniella)(Hongia)(Kribbella)(Marmoricola)(Micropruina)(Nocardioides)(Pimelobacter)(Propionicicella)(Propionicimonas)丙酸桿菌科(Propionibacteriaceae)河口微菌屬(Aestuariimicrobium)(Arachnia)(Brooklawnia)(Granulicoccus)江氏菌屬(Jiangella)(Luteococcus)(Microlunatus)丙酸桿菌屬(Propionibacterium)(Propioniferax)(Propionimicrobium)(Tessaracocccus)(Pseudonocardineae)(Actinosynnemataceae)(Actinokineospora)(Actinosynnema)(Lechevalieria)(Lentzea)(Saccharothrix)(Umezawaea)(Pseudonocardiaceae)(Actinoalloteichus)(Actinobispora)(Amycolata)(Amycolatopsis)(Crossiella)(Faenia)(Goodfellowia)(Kibdelosporangium)(Kutzneria)(Prauserella)(Pseudoamycolata)(Pseudonocardia)(Saccharomonospora)(Saccharopolyspora)(Streptoalloteichus)(Thermobispora)(Thermocrispum)鏈霉菌亞目(Streptomycineae)鏈霉菌科(Sterptomycetaceae)(Actinopycnidium)(Actinosporangium)(Chainia)(Elytrosporangium)北里菌屬(Kitasatoa)北里孢菌屬(Kitasatospora)(Microellobosporia)(Streptacidiphilus)鏈霉菌屬(Streptomyces)(Streptoverticillium)(Streptosporangineae)(Nocardiopsaceae)(Nocardiopsis)(Streptomonospora)(Thermobifida)(Streptosporangiaceae)(Acrocarpospora)(Herbidospora)(Microbispora)(Microtetraspora)(Nonomuraea)(Planobispora)(Planomonospora)(Planotetraspora)(Sphaerisporangium)(Streptosporangium)(Thermopolyspora)(Thermomonosporaceae)珊瑚狀放線菌屬(Actinocorallia)(Actinomadura)(Excellospora)(Spirillospora)(Thermomonospora)雙歧桿菌目(Bifidobacteriales)雙歧桿菌科(Bifidobacteriaceae)(Aeriscardovia)(Alloscardovia)雙歧桿菌屬(Bifidobacterium)(Gardnerella)(Metascardovia)(Parascardovia)(Scardovia)科未定(Coriobacteridae)(Coriobacteriales)(Coriobacterineae)(Coriobacteriaceae)奇異菌屬(Atopobium)(Collinsella)(Coriobacterium)(Cryptobacterium)(Denitrobacterium)(Eggerthella)(Olsenella)(Slackia)2.4紅色桿菌亞綱(Rubrobacteridae)

紅色桿菌目(Rubrobacterales)紅色桿菌亞目(Rubrobacterineae)(Conexibacteraceae)(Conexibacter)(Patulibacteraceae)(Patulibacter)紅色桿菌科(Rubrobacteraceae)紅色桿菌屬(Rubrobacter)(Solirubrobacteraceae)(Solirubrobacter)(Thermoleophilaceae)(Thermoleophilum)2.5球形桿菌亞綱(Sphaerobacteridae)

球形桿菌目(Sphaerobacterales)球形桿菌亞目(Sphaerobacterineae)球形桿菌科(Sphaerobacteraceae)球形桿菌屬(Sphaerobacter)3產(chǎn)水菌門(Aquificae)3.1產(chǎn)水菌綱(Aquificae)產(chǎn)水菌目(Aquificales)產(chǎn)水菌科(Aquificaceae)產(chǎn)水菌屬(Aquifex)(Calderobacterium)(Hydrogenivirga)(Hydrogenobacter)(Hydrogenobaculum)(Thermocrinis)除硫桿菌科(Desulfurobacteriaceae)(Balnearium)除硫桿菌屬(Desulfurobacterium)熱弧菌屬(Thermovibrio)(Hydrogenothermaceae)(Hydrogenothermus)(Persephonella)(Sulfurihydrogenibium)4擬桿菌門(Bacteroidetes)4.1擬桿菌綱(Bacteroidetes)擬桿菌目(Bacteroidales)擬桿菌科(Bacteroidaceae)(Acetomicrobium)(Anaerophaga)(Anaerorhabdus)擬桿菌屬(Bacteroides)(Pontibacter)紫單胞菌科(Porphyromonadaceae)(Barnesiella)(Capsularis)(Dysgonomonas)(Hallella)(Odoribacter)(Oribaculum)(Paludibacter)(Parabacteroides)紫單胞菌屬(Porphyromonas)（多譯作“卟啉單胞菌”，但porphyro-應(yīng)來源于希臘語“紫色”）(Proteiniphilum)(Tannerella)(Xylanibacter)普雷沃氏菌科(Prevotellaceae)普雷沃氏菌屬(Prevotella)

（或譯作“普氏菌”）理研菌科(Rikenellaceae)(Alistipes)(Alkaliflexus)(Marinilabilia)(Petrimonas)理研菌屬(Rikenella)

（注：Riken是日語“理化學(xué)研究所”簡稱）科未定

(Acetofilamentum)(Acetothermus)4.2黃桿菌綱(Flavobacteria)黃桿菌目(Flavobacteriales)蟑螂桿狀體科(Blattabacteriaceae)蟑螂桿狀體屬(Blattabacterium)(Cryomorphaceae)(Algoriphagus)(Brumimicrobium)(Crocinitomix)(Cryomorpha)(Fluviicola)李時珍菌屬(Lishizhenia)(Owenweeksia)黃桿菌科(Flavobacteriaceae)(Actibacter)(Aequorivita)(Algibacter)(Aquimarina)(Arenibacter)伯杰菌屬(Bergeyella)(Bizionia)碳酸噬胞菌屬(Capnocytophaga)

（注：多譯作“二氧化碳噬纖維菌屬”）噬纖維素菌屬(Cellulophaga)金黃桿菌屬(Chryseobacterium)(Cloacibacterium)(Coenonia)(Costertonia)(Croceibacter)獨(dú)島菌屬(Dokdonia)東海菌屬(Donghaeana)(Elizabethkingia)(Empedobacter)(Epilithonimonas)(Flaviramulus)黃桿菌屬(Flavobacterium)(Formosa)泥灘桿菌屬(Gaetbulibacter)泥灘微菌屬(Gaetbulimicrobium)(Galbibacter)(Gelidibacter)(Gillisia)(Gilvibacter)革蘭菌屬(Gramella)(Kaistella)(Kordia)(Krokinobacter)(Lacinutrix)列文虎克菌屬(Leeuwenhoekiella)(Lutibacter)(Maribacter)(Mariniflexile)(Marixanthomonas)(Mesonia)(Muricauda)(Myroides)(Nonlabens)(Olleya)(Ornithobacterium)(Persicivirga)(Pibocella)極地桿菌屬(Polaribacter)冷彎菌屬(Psychroflexus)(Psychroserpens)(Riemerella)(Robiginitalea)(Salegentibacter)(Sandarakinotalea)(Sediminibacter)(Sediminicola)世宗菌屬(Sejongia)(Stanierella)(Stenothermobacter)(Subsaxibacter)(Subsaximicrobium)(Tamlana)(Tenacibaculum)(Ulvibacter)(Vitellibacter)(Wautersiella)(Weeksella)(Winogradskyella)麗水菌屬(Yeosuana)(Zeaxanthinibacter)周氏菌屬(Zhouia)(Zobellia)王祖農(nóng)菌屬(Zunongwangia)4.3鞘脂桿菌綱(Sphingobacteria)鞘脂桿菌目(Sphingobacteriales)泉發(fā)菌科(Crenotrichaceae)(Balneola)(Chitinophaga)泉發(fā)菌屬(Crenothrix)(Rhodothermus)(Salinibacter)(Terrimonas)(Toxothrix)(Flammeovirgaceae)(Flammeovirga)(Flexithrix)(Perexilibacter)(Persicobacter)(Rapidithrix)(Sediminitomix)(Thermonema)屈撓桿菌科(Flexibacteraceae)(Adhaeribacter)(Aquiflexum)(Arcicella)(Belliella)(Chimaereicella)(Cyclobacterium)噬胞菌屬(Cytophaga)

（注：目前多稱此屬為“噬纖維菌屬”，此處依拉丁文）(Dyadobacter)(Echinicola)(Effluviibacter)(Emticicia)(Fabibacter)(Flectobacillus)屈撓桿菌屬(Flexibacter)(Hongiella)(Hymenobacter)(Larkinella)(Leadbetterella)(Marinicola)(Meniscus)(Microscilla)(Niastella)(Persicitalea)(Reichenbachiella)(Rhodonellum)(Roseivirga)(Runella)(Spirosoma)生孢噬胞菌屬(Sporocytophaga)腐螺旋菌科(Saprospiraceae)(Aureispira)(Haliscomenobacter)(Lewinella)腐螺旋菌屬(Saprospira)鞘脂桿菌科(Sphingobacteriaceae)(Mucilaginibacter)(Olivibacter)(Parapedobacter)(Pedobacter)(Pseudosphingobacterium)鞘脂桿菌屬(Sphingobacterium)科未定

(Niabella)4.4綱未定(Flavisolibacter)(Fulvivirga)(Prolixibacter)(Segetibacter)5衣原體門(Chlamydiae)5.1衣原體綱(Chlamydiae)衣原體目(Chlamydiales)衣原體科(Chlamydiaceae)衣原體屬(Chlamydia)嗜衣體屬(Chlamydophila)副衣原體屬(Parachlamydiaceae)新衣原體屬(Neochlamydia)副衣原體屬(Parachlamydia)芯卡體科(Simkaniaceae)芯卡體屬(Simkania)（注：人名縮寫簡稱，此處用音譯）棍衣原體屬(Rhabdochlamydia)*華診體科(Waddliaceae)華診體屬(Waddlia)

（注：WADDL為“華盛頓動物病診斷實(shí)驗室”縮寫）6綠菌門(Chlorobi)6.1綠菌綱(Chlorobia)綠菌目(Chlorobiales)綠菌科(Chlorobiaceae)臂綠菌屬(Ancalochloris)綠棒菌屬(Chlorobaculum)綠菌屬(Chlorobium)綠爬菌屬(Chloroherpeton)暗網(wǎng)菌屬(Pelodictyon)突柄綠菌屬(Prosthecochloris)7綠彎菌門(Chloroflexi)7.1厭氧繩菌綱(Anaerolineae)厭氧繩菌目(Anaerolineales)厭氧繩菌科(Anaerolineaceae)厭氧繩菌屬(Anaerolinea)(Bellilinea)纖繩菌屬(Leptolinea)(Levilinea)長繩菌屬(Longilinea)7.2暖繩菌綱(Caldilineae)暖繩菌目(Caldilineales)暖繩菌科(Caldilineaceae)暖繩菌屬(Caldilinea)7.3綠彎菌綱(Chloroflexi)綠彎菌目(Chloroflexales)綠彎菌科(Chloroflexaceae)綠彎菌屬(Chloroflexus)綠線菌屬(Chloronema)太陽發(fā)菌屬(Heliothrix)玫瑰彎菌屬(Roseiflexus)顫綠菌科(Oscillochloridaceae)顫綠菌屬(Oscillochloris)爬管菌目(Herpetosiphonales)爬管菌科(Herpetosiphonaceae)爬管菌屬(Herpetosiphon)8產(chǎn)金菌門(Chrysiogenetes)8.1產(chǎn)金菌綱(Chrysiogenetes)產(chǎn)金菌目(Chrysiogenales)產(chǎn)金菌科(Chrysiogenaceae)產(chǎn)金菌屬(Chrysiogenes)9藍(lán)藻門(Cyanobacteria)9.1藍(lán)藻綱(Cyanobacteria)注：目前有三套藍(lán)藻分類系統(tǒng)，分別為NCBI、Bergey's手冊及Cavalier-Smith（2002年，僅分至目)。本文采用的Bergey's手冊中藍(lán)藻內(nèi)的分類，不論目科屬，均為形態(tài)分類，其中目在手冊中為“Subsection”而非“Order”，目、科均未列名稱，此處目名稱為習(xí)慣用法，科不列名稱；屬除Halospirulina和Trichodesmium之外均標(biāo)為“形態(tài)屬”(FormGenus)而非“屬”(Genus)。表中在形態(tài)分類后面加注'@'。預(yù)料不久的將來整個藍(lán)藻分類系統(tǒng)將會大調(diào)，與其它細(xì)菌統(tǒng)一為系統(tǒng)發(fā)育分類系統(tǒng)。此外，因漢語“藻”字可指多種水生光合產(chǎn)氧生物，包括多支互不相關(guān)的原生生物甚至高等水生植物，此處仍用按照“藻”翻譯藍(lán)藻門(Cyanobacteria)細(xì)菌。但也可依據(jù)個人理念將名稱定為“某菌”或“某藍(lán)菌”，但應(yīng)注意避免與其余細(xì)菌重名。第一目（色球藻目(Chroococcales)）@第一科@管孢藻屬(Chamaesiphon)@色球藻屬(Chroococcus)@(Cyanobacterium)@(Cyanobium)@(Cyanothece)@藍(lán)纖維藻屬(Dactylococcopsis)@(Gloeobacter)@黏球藻屬(Gloeocapsa)@黏桿藻屬(Gloeothece)@微囊藻屬(Microcystis)@原綠球藻屬(Prochlorococcus)@原綠藻屬(Prochloron)@聚球藻屬(Synechococcus)@集胞藻屬(Synechocystis)@第二目（寬球藻目(Pleurocapsales)）@第一科@(Cyanocystis)@(Dermocarpella)@(Stanieria)@異球藻屬(Xenococcus)@第二科@擬色球藻屬(Chroococcidiopsis)@黏囊藻屬(Myxosarcina)@寬球藻屬(Pleurocapsa)@第三目（顫藻目(Oscillatoriales)）@第一科@節(jié)螺藻屬(Arthrospira)@

（包含商業(yè)化螺旋藻）(Borzia)@(Crinalium)@(Geitlerinema)@鹽螺旋藻屬(Halospirulina)(Leptolyngbya)@(Limnothrix)@鞘絲藻屬(Lyngbya)@微鞘藻屬(Microcoleus)@顫藻屬(Oscillatoria)@(Planktothrix)@原綠發(fā)藻屬(Prochlorothrix)@(Pseudanabaena)@螺旋藻屬(Spirulina)@(Starria)@束藻屬(Symploca)@束毛藻屬(Trichodesmium)(Tychonema)@第四目（念珠藻目(Nostocales)）@第一科@魚腥藻屬(Anabaena)@項圈藻屬(Anabaenopsis)@束絲藻屬(Aphanizomenon)@(Cyanospira)@(Cylindrospermopsis)@筒孢藻屬(Cylindrospermum)@節(jié)球藻屬(Nodularia)@念珠藻屬(Nostoc)@偽枝藻屬(Scytonema)@第二科@眉藻屬(Calothrix)@膠須藻屬(Rivularia)@單歧藻屬(Tolypothrix)@第五目（真枝藻目(Stigonematales)）@第一科@(Chlorogloeopsis)@費(fèi)氏藻屬(Fischerella)@(Geitleria)@(Iyengariella)@擬念珠藻屬(Nostochopsis)@真枝藻屬(Stigonema)@10脫鐵桿菌門(Deferribacteres)10.1脫鐵桿菌綱(Deferribacteres)脫鐵桿菌目(Deferribacterales)脫鐵桿菌科(Deferribacteraceae)脫鐵桿菌屬(Deferribacter)(Denitrovibrio)(Flexistipes)地弧菌屬(Geovibrio)(Mucispirillum)科未定

暖發(fā)菌屬(Caldithrix)(Synergistes)11異常球菌-棲熱菌門(Deinococcus-Thermus)11.1異常球菌綱(Deinococci)異常球菌目(Deinococcales)異常球菌科(Deinococcaceae)異常桿菌屬(Deinobacter)異常球菌屬(Deinococcus)特呂珀菌科(Trueperaceae)特呂珀菌屬(Truepera)棲熱菌目(Thermales)棲熱菌科(Thermaceae)海棲熱菌屬(Marinithermus)亞棲熱菌屬(Meiothermus)海洋棲熱菌屬(Oceanithermus)棲熱菌屬(Thermus)火山棲熱菌屬(Vulcanithermus)12網(wǎng)團(tuán)菌門(Dictyoglomi)12.1網(wǎng)團(tuán)菌綱(Dictyoglomi)網(wǎng)團(tuán)菌目(Dictyoglomales)網(wǎng)團(tuán)菌科(Dictyoglomaceae)網(wǎng)團(tuán)菌屬(Dictyoglomus)13纖維桿菌門(Fibrobacteres)13.1纖維桿菌綱(Fibrobacteres)纖維桿菌目(Fibrobacterales)纖維桿菌科(Fibrobacteraceae)纖維桿菌屬(Fibrobacter)14厚壁菌門(Firmicutes)（低G+C革蘭氏陽性菌）14.1芽孢桿菌綱(Bacilli)芽孢桿菌目(Bacillales)(Alicyclobacillaceae)(Alicyclobacillus)(Pasteuria)(Sulfobacillus)芽孢桿菌科(Bacillaceae)(Alkalibacillus)(Amphibacillus)(Anoxybacillus)芽孢桿菌屬(Bacillus)(Caldalkalibacillus)(Cerasibacillus)(Exiguobacterium)(Filobacillus)(Geobacillus)(Gracilibacillus)(Halalkalibacillus)(Halobacillus)(Haloactibacillus)(Jeotgalibacillus)(Lentibacillus)(Lysinibacillus)(Marinibacillus)(Oceanobacillus)(Ornithinibacillus)(Paraliobacillus)(Paucisalibacillus)(Pelagibacillus)(Piscibacillus)(Pontibacillus)糖球菌屬(Saccharococcus)(Salibacillus)(Salimicrobium)(Salinibacillus)(Salirhabdus)(Sasuginibacillus)(Tenuibacillus)(Terribacillus)(Thalassobacillus)(Ureibacillus)(Virgibacillus)(Vulcanibacillus)(Caryophanaceae)(Caryophanon)李斯特氏菌科(Listeriaceae)環(huán)絲菌屬(Brochothrix)李斯特氏菌屬(Listeria)(Paenibacillaceae)(Ammoniphilus)(Aneurinibacillus)(Brevibacillus)(Cohnella)(Oxalophagus)(Paenibacillus)(Thermicanus)(Thermobacillus)(Planococcaceae)(Filibacter)(Kurthia)(Planococcus)(Planomicrobium)(Sporosarcina)芽孢乳桿菌科(Sporolactobacillaceae)(Marinococcus)(Sinococcus)芽孢乳桿菌屬(Sporolactobacillus)(Tuberibacillus)葡萄球菌科(Staphylococcaceae)孿生菌屬(Gemella)(Joetgalicoccus)(Macrococcus)(Salinicoccus)葡萄球菌屬(Staphylococcus)(Thermoactinomycetaceae)(Laceyella)(Mechercharimyces)(Planifilum)(Seinonella)(Shimazuella)(Thermoactinomyces)(Thermoflavimicrobium)(Turicibacteraceae)(Turicibacter)科未定(Pullulanibacillus)乳桿菌目(Lactobacillales)氣球菌科(Aerococcaceae)(Abiotrophia)氣球菌屬(Aerococcus)(Dolosicoccus)(Eremococcus)(Facklamia)(Globicatella)(Ignavigranum)肉桿菌科(Carnobacteriaceae)(Agitococcus)(Alkalibacterium)(Allofustis)(Alloiococcus)(Atopococcus)(Atopostipes)肉桿菌屬(Carnobacterium)德庫菌屬(Desemzia)

（DSMZ，德國微生物與細(xì)胞保藏中心）(Dolosigranulum)(Granulicatella)(Isobaculum)(Lactosphaera)(Marinilactibacillus)(Trichococcus)腸球菌科(Enterococcaceae)(Atopobacter)(Catellicoccus)腸球菌屬(Enterococcus)(Melissococcus)(Pilibacter)(Tetragenococcus)漫游球菌屬(Vagococcus)乳桿菌科(Lactobacillaceae)乳桿菌屬(Lactobacillus)(Paralactobacillus)片球菌屬(Pediococcus)明串珠菌科(Leuconostocaceae)明串珠菌屬(Leuconostoc)(Oenococcus)(Weissella)顫螺旋菌科(Oscillospiraceae)顫桿菌克(Oscillibacter)顫螺旋菌屬(Oscillospira)鏈球菌科(Streptococcaceae)乳球菌屬(Lactococcus)(Lactovum)鏈球菌屬(Streptococcus)科未定

(Acetoanaerobium)互營球菌屬(Syntrophococcus)14.2梭菌綱(Clostridia)梭菌目(Clostridiales)梭菌科(Clostridiaceae)(Acetanaerobacterium)(Acetivibrio)(Acidaminobacter)(Alkaliphilus)(Anaerobacter)(Anaerotruncus)(Anoxynatronum)(Bryantella)(Caloramator)(Caloranaerobacter)(Caminicella)(Clostridiisalibacter)梭菌屬(Clostridium)(Coprobacillus)(Dorea)(Ethanoligenens)(Faecalibacterium)(Fastidiosipila)(Garciella)(Gracilibacter)(Guggenheimella)(Hespellia)(Natronincola)(Oxobacter)(Parasporobacterium)八疊球菌屬(Sarcina)(Soehngenia)(Sporacetigenium)(Sporobacter)(Subdoligranulum)(Tepidibacter)(Tepidimicrobium)(Thermobrachium)(Thermohalobacter)(Tindallia)優(yōu)桿菌科(Eubacteriaceae)醋桿菌屬(Acetobacterium)(Alkalibacter)(Anaerofustis)(Anaerovorax)優(yōu)桿菌屬(Eubacterium)(Mogibacterium)(Pseudoramibacter)太陽桿菌科(Heliobacteriaceae)太陽桿狀菌屬(Heliobacillus)太陽桿菌屬(Heliobacterium)嗜日菌屬(Heliophilum)(Heliorestis)毛螺旋菌科(Lachnospiraceae)(Acetitomaculum)(Anaerofilum)(Anaerostipes)丁酸弧菌屬(Butyrivibrio)(Catenibacterium)(Catonella)糞球菌屬(Coprococcus)(Johnsonella)(Lachnobacterium)毛螺旋菌屬(Lachnospira)(Moryella)(Oribacterium)假丁酸弧菌屬(Pseudobutyrivibrio)(Roseburia)瘤胃球菌屬(Ruminococcus)(Shuttleworthia)(Sporobacterium)消化球菌科(Peptococcaceae)(Carboxydothermus)(Cryptanaerobacter)(Dehalobacter)(Desulfitobacterium)(Desulfonispora)(Desulfosporosinus)(Desulfotomaculum)(Pelotomaculum)消化球菌屬(Peptococcus)(Syntrophobotulus)(Thermincola)(Thermoterrabacterium)消化鏈球菌科(Peptostreptococcaceae)(Anaerococcus)(Filifactor)(Finegoldia)(Fusibacter)(Gallicola)(Helcococcus)微單胞菌屬(Parvimonas)

（原名Micromonas）(Peptoniphilus)消化鏈球菌屬(Peptostreptococcus)(Sedimentibacter)(Sporanaerobacter)(Tissierella)互營單胞菌科(Syntrophomonadaceae)(Aminiphilus)(Aminobacterium)(Aminomonas)(Anaerobaculum)(Anaerobranca)(Caldicellulosiruptor)(Carboxydocella)(Dethiosulfovibrio)(Pelospora)互營單胞菌屬(Syntrophomonas)(Syntrophospora)互營熱菌屬(Syntrophothermus)(Thermaerobacter)(Thermanaerovibrio)(Thermohydrogenium)(Thermosyntropha)(Thermovirga)韋榮氏菌科(Veillonellaceae)(Acetonema)氨基酸球菌屬(Acidaminococcus)(Allisonella)(Anaeroarcus)(Anaeroglobus)(Anaeromusa)(Anaerosinus)(Anaerovibrio)(Centipeda)(Dendrosporobacter)(Dialister)(Megamonas)(Megasphaera)光岡菌屬(Mitsuokella)(Papillibacter)(Pectinatus)(Pelosinus)(Phascolarctobacterium)(Propionispira)(Propionispora)(Quinella)(Schwartzia)新月形單胞菌屬(Selenomonas)鼠孢菌屬(Sporomusa)(Sporotalea)(Succiniclasticum)(Succinispira)(Thermosinus)韋榮氏菌屬(Veillonella)(Zymophilus)科未定

(Anaerovirgula)(Dethiosulfatibacter)(Geosporobacter)(Howardella)(Proteiniborus)鹽厭氧菌目(Halanaerobiales)鹽厭氧菌科(Halanaerobiaceae)鹽厭氧菌屬(Halanaerobium)(Halocella)(Haloincola)(Halothermothrix)鹽擬桿菌科(Halobacteroidaceae)(Acetohalobium)(Halanaerobacter)鹽擬桿菌屬(Halobacteroides)(Halonatronum)(Natroniella)(Orenia)(Selenihalanaerobacter)(Sporohalobacter)鹽堿厭氧菌目(Natranaerobiales)鹽堿厭氧菌科(Natranaerobiaceae)鹽堿厭氧菌屬(Natranaerobius)熱厭氧桿菌目(Thermoanaerobacteriales)熱厭氧桿菌科(Thermoanaerobacteriaceae)(Acetogenium)(Ammonifex)(Caldanaerobacter)(Carboxydibrachium)(Coprothermobacter)(Desulfovirgula)(Gelria)(Mahella)(Moorella)(Sporotomaculum)(Tepidanaerobacter)(Thermacetogenium)熱厭氧單胞菌屬(Thermanaeromonas)(Thermoanaerobacter)(Thermoanaerobium)(Thermoanaerobacterium)熱厭氧菌屬(Thermoanaerobium)(Thermobacteroides)(Thermosediminibacter)(Thermovenabulum)熱脫硫菌科(Thermodesulfobiaceae)熱脫硫菌屬(Thermodesulfobium)目未定

(Desulfitibacter)(Symbiobacterium)14.3熱石桿菌綱(Thermolithobacteria)熱石菌目(Thermolithobacterales)

熱石桿菌科(Thermolithobacteraceae)

熱石桿菌屬(Thermolithobacter)15梭桿菌門(Fusobacteria)15.1梭桿菌綱(Fusobacteria)梭桿菌目(Fuobacteriales)

梭桿菌科(Fusobacteriaceae)

梭桿菌屬(Fusobacterium)(Ilyobcter)纖毛菌屬(Letotrichia)(Propiongenium)(Sebaldel)(Sneathia)(reptobacils)科未定

(Cetobacteium)16芽單胞菌門(Gemmatimonadetes)16.1芽單胞菌綱(Gemmatimonadetes)芽單胞菌目(Gemmatimonadals)芽單胞菌科(Gemmatimonadaceae)芽單胞菌屬(Gemmatimonas)17黏膠球形菌門(Lentisphaerae)17.1黏膠球形菌綱(Lentisphaerae)黏膠球形菌目(Lentisphaerale)黏膠球形菌科(Lentisphaeraceae)黏膠球形菌屬(Lentisphaera)食物谷菌目(Victivallales)食物谷菌科(Victivallaceae)食物谷菌屬(Victivallis)18硝化螺旋菌門(Nitrospirae)18.1硝化螺旋菌綱(Nitrospira)硝化螺旋菌目(Nitrospirales)硝化螺旋菌科(Nitrospiraceae)鉤端螺菌屬(Leptospirillum)硝化螺旋菌屬(Nitrospira)磁桿菌屬(Magnetobacterium)*熱脫硫弧菌屬(Thermodesulfovibrio)19浮霉菌門(Planctomycetes)19.1浮霉菌綱(Planctomycetacia)浮霉菌目(Planctomycetales)浮霉菌科(Planctomycetaceae)(Blastopirellula)(Gemmata)(Isosphaera)小梨形菌屬(Pirellula)浮霉菌屬(Planctomycetes)紅小梨形菌屬(Rhodopirellula)(Schlesneria)科未定

（均未能分離培養(yǎng)，見厭氧氨氧化菌）(Brocadia)*(Kuenenia)*(Scalindua)*20海綿桿菌門(Poribacteria)*21變形菌門(Proteobacteria)21.1α-變形菌綱(Alphaproteobacteria)柄桿菌目(Caulobacterales)柄桿菌科(Caulobacteraceae)(Asticcacaulis)短波單胞菌屬(Brevundimonas)柄桿菌屬(Caulobacter)苯基桿菌屬(Phenylobacterium)(Kordiimonadales)(Kordiimonadaceae)(Kordiimonas)短小盒菌目(Parvularculales)短小盒菌科(Parvularculaceae)短小盒菌屬(Parvularcula)根瘤菌目(Rhizobiales)橙單胞菌科(Aurantimonadaceae)橙單胞菌屬(Aurantimonas)(Fulvimarina)(Martelella)巴通體科(Bartonellaceae)巴通體屬(Bartonella)(Grahamella)(Rochalimaea)拜葉林克氏菌科(Beijerinckiaceae)拜葉林克氏菌屬(Beijerinckia)(Chelatococcus)(Methylocapsa)(Methylocella)慢生根瘤菌科(Bradyrhizobiaceae)阿菲波菌屬(Afipia)農(nóng)單胞菌屬(Agromonas)(Balneimonas)芽生桿菌屬(Blastobacter)(Bosea)慢生根瘤菌屬(Bradyrhizobium)硝化桿菌屬(Nitrobacter)(Oligotropha)(Rhodoblastus)紅假單胞菌屬(Rhodopseudomonas)布魯氏菌科(Brucellaceae)布魯氏菌屬(Brucella)(Crabtreella)(Daeguia)支動菌屬(Mycoplana)蒼白桿菌屬(Ochrobactrum)(Pseudochrobactrum)黏著桿菌科(Cohaesibacteraceae)黏著桿菌屬(Cohaesibacter)生絲微菌科(Hyphomicrobiaceae)(Ancalomicrobium)(Angulomicrobium)水桿菌屬(Aquabacter)芽綠菌屬(Blastochloris)(Devosia)(Dichotomicrobium)(Filomicrobium)(Gemmiger)生絲微菌屬(Hyphomicrobium)(Methylorhabdus)土微菌屬(Pedomicrobium)突柄微菌屬(Prosthecomicrobium)紅微菌屬(Rhodomicrobium)(Rhodoplanes)(Seliberia)甲基桿菌科(Methylobacteriaceae)(Meganema)甲基桿菌屬(Methylobacterium)(Microvirga)(Protomonas)甲基孢囊菌科(Methylocystaceae)(Albibacter)甲基孢囊菌屬(Methylocystis)(Methylopila)(Methylosinus)(Pleomorphomonas)(Terasakiella)葉桿菌科(Phyllobacteriaceae)(Aminobacter)(Aquamicrobium)(Defluvibacter)(Hoeflea)中慢生根瘤菌屬(Mesorhizobium)(Nitratireductor)葉桿菌屬(Phyllobacterium)(Pseudaminobacter)根瘤菌科(Rhizobiaceae)土壤桿菌屬(Agrobacterium)異根瘤菌屬(Allorhizobium)嗜碳菌屬(Carbophilus)螯合桿菌屬(Chelatobacter)劍菌屬(Ensifer)(Kaistia)韌皮桿菌屬(Liberibacter)*根瘤菌屬(Rhizobium)中華根瘤菌屬(Sinorhizobium)紅菌科(Rhodobiaceae)(Anderseniella)(Parvibaculum)紅菌屬(Rhodobium)玫瑰螺菌屬(Roseospirillum)黃色桿菌科(Xanthobacteraceae)(Ancylobacter)(Azorhizobium)(Labrys)(Pseudolabrys)(Starkeya)黃色桿菌屬(Xanthobacter)科未定(Sneathiella)紅桿菌目(Rhodobacterales)生絲單胞菌科(Hyphomonadaceae)(Hirschia)生絲單胞菌屬(Hyphomonas)(Maricaulis)(Oceanicaulis)(Robiginitomaculum)紅桿菌科(Rhodobacteraceae)(Ahrensia)(Albidovulum)(Amaricoccus)(Antarctobacter)(Catellibacterium)(Citreicella)(Citreimonas)(Dinoroseobacter)(Donghicola)(Gemmobacter)(Haematobacter)(Jannaschia)(Ketogulonicigenium)(Labrenzia)(Leisingera)(Loktanella)(Maribius)(Marinovum)(Maritimibacter)(Methylarcula)(Nereida)(Nesiotobacter)(Oceanibulbus)海洋生菌屬(Oceanicola)(Octadecabacter)(Palleronia)(Pannonibacter)副球菌屬(Paracoccus)(Pelagibaca)(Phaeobacter)假紅桿菌屬(Pseudorhodobacter)(Pseudovibrio)(Rhodobaca)紅桿菌屬(Rhodobacter)(Rhodothalassium)小紅卵菌屬(Rhodovulum)(Roseibacterium)(Roseibium)(Roseicyclus)(Roseinatronobacter)(Roseisalinus)(Roseivivax)玫瑰桿菌屬(Roseobacter)(Roseovarius)(Rubellimicrobium)(Rubrimonas)(Ruegeria)(Sagittula)(Salipiger)(Shimia)(Silicibacter)(Staleya)(Stappia)(Sulfitobacter)(Tateyamaria)(Thalassobacter)(Thalassobius)(Thalassococcus)(Thioclava)硫球形菌屬(Thiosphaera)(Wenxinia)木洞菌屬(Woodsholea)楊氏菌屬(Yangia)紅螺菌目(Rhodospirillales)醋桿菌科(Acetobacteraceae)醋桿菌屬(Acetobacter)(Acidicaldus)嗜酸菌屬(Acidiphilium)酸球形菌屬(Acidisphaera)酸胞菌屬(Acidocella)酸單胞菌屬(Acidomonas)(Asaia)(Belnapia)脆弱球菌屬(Craurococcus)葡糖醋桿菌屬(Gluconacetobacter)葡糖桿菌屬(Gluconobacter)(Granulibacter)(Kozakia)鼠球菌屬(Muricoccus)(Neoasaia)(Paracraurococcus)紅球狀菌屬(Rhodopila)(Rhodovarius)玫瑰球菌屬(Roseococcus)玫瑰單胞菌屬(Roseomonas)(Rubritepida)(Saccharibacter)(Stella)(Swaminathania)(Teichococcus)(Zavarzinia)紅螺菌科(Rhodospirillaceae)固氮螺菌屬(Azospirillum)(Conglomeromonas)(Defluviicoccus)(Inquilinus)磁螺菌屬(Magnetospirillum)(Phaeospirillum)(Rhodocista)紅螺旋菌屬(Rhodospira)紅螺菌屬(Rhodospirillum)紅弧菌屬(Rhodovibrio)玫瑰螺旋菌屬(Roseospira)(Skermanella)(Telmatospirillum)(Thalassospira)(Tistrella)立克次體目(Rickettsiales)無形體科(Anaplasmataceae)(Aegyptianella)無形體屬(Anaplasma)(Cowdria)埃里希體屬(Ehrlichia)新立克次體屬(Neorickettsia)沃巴赫體屬(Wolbachia)(Xenohaliotis)*全孢菌科(Holosporaceae)全孢菌屬(Holospora)立克次體科(Rickettsiaceae)(Orientia)立克次體屬(Rickettsia)SAR11群*遠(yuǎn)洋桿菌屬(Pelagibacter)*科未定(Lyticum)(Odyssella)*(Pseudocaedibacter)(Symbiotes)(Tectibacter)鞘脂單胞菌目(Sphingomonadales)赤桿菌科(Erythrobacteraceae)(Altererythrobacter)赤桿菌屬(Erythrobacter)赤微菌屬(Erythromicrobium)紫桿菌屬(Porphyrobacter)鞘脂單胞菌科(Sphingomonadaceae)芽單胞菌屬(Blastomonas)檸檬酸微菌屬(Citromicrobium)*赤單胞菌屬(Erythromonas)新鞘脂菌屬(Novosphingobium)(Sandaracinobacter)(Sandaracinorhabdus）鞘脂菌屬(Sphingobium)鞘脂單胞菌屬(Sphingomonas)(Sphingopyxis)(Sphingosinicella)發(fā)酵單胞菌屬(Zymomonas)目未定(Albimonas)(Caenispirillum)(Insolitispirillum)(Novispirillum)21.2

β-變形菌綱(Betaproteobacteria)伯克氏菌目(Burkholderiales)產(chǎn)堿菌科(Alcaligenaceae)無色桿菌屬(Achromobacter)(Advenella)產(chǎn)堿菌屬(Alcaligenes)(Azohydromonas)博德氏菌屬(Bordetella)(Brackiella)(Castellaniela)德克斯氏菌屬(Derxia)(Kerstersia)(Oligella)(Pelistega)(Pigmentiphaga)(Pusillimonas)(Sutterella)(Taylorella)(Tetrathiobacter)伯克氏菌科(Burkholderiaceae)伯克氏菌屬(Burkholderia)(Chitinimonas)(Cupriavidus)(Lautropia)(Limnobacter)(Pandoraea)(Paucimonas)(Polynucleobacter)青枯菌屬(Ralstonia)(Thermothrix)沃特氏菌屬(Wautersia)叢毛單胞菌科(Comamonadaceae)食酸菌屬(Acidovorax)(Alcycliphilus)(Brachymonas)(Caenibacterium)(Caldimonas)叢毛單胞菌屬(Comamonas)(Curvibacter)代爾夫特菌屬(Delftia)(Diaphorobacter)(Giesbergeria)噬氫菌屬(Hydrogenophaga)(Hylemonella)(Lampropedia)(Macromonas)(Malikia)(Ottowia)(Pelomonas)(Polaromonas)(Ramlibacter)紅育菌屬(Rhodoferax)(Schlegelella)(Simplicispira)貪食菌屬(Variovorax)(Xenophilus)草酸桿菌科(Oxalobacteraceae)(Collimonas)(Duganella)(Herbaspirillum)(Herminiimonas)(Janthinobacterium)(Massilia)(Naxibacter)(Oxalicibacterium)草酸桿菌屬(Oxalobacter)(Telluria)(Undibacterium)科未定(Aquabacterium)(Aquincola)(Ideonella)纖發(fā)菌屬(Leptothrix)(Methylibium(Mitsuaria)(Paucibacter)(Roseateles)紅長命菌屬(Rubrivivax)(Sphaerotilus)(Tepidicella)(Tepidimonas)(Thiobacter)硫單胞菌屬(Thiomonas)(Xylophilus)嗜氫菌目(Hydrogenophilales)嗜氫菌科(Hydrogenophilaceae)嗜氫菌屬(Hydrogenophilus)(Petrobacter)(Tepidiphilus)硫桿狀菌屬(Thiobacillus)嗜甲基菌目(Methylophilales)嗜甲基菌科(Methylophilaceae)(Methylobacillus)嗜甲基菌屬(Methylophilus)(Methylotenera)(Methylovorus)奈瑟菌目(Neisseriales)奈瑟菌科(Neisseriaceae)(Alysiella)(Andreprevotia)水螺菌屬(Aquaspirillum)(Aquitalea)(Bergeriella)(Chitinibacter)(Chitinilyticum)色桿菌屬(Chromobacterium)(Conchiformibius)(Deefgea)艾肯氏菌屬(Eikenella)(Formivibrio)(Gulbenkiania)(Iodobacter)金氏菌屬(Kingella)鷗桿菌屬(Laribacter)(Leeia)(Microvirgula)(Morococcus)奈瑟菌屬(Neisseria)(Paludibacterium)(Prolinoborus)(Silvimonas)(Simonsiella)(Uruburuella)(Vitreoscilla)(Vogesella)亞硝化單胞菌目(Nitrosomonadales)嘉利翁氏菌科(Gallionellaceae)嘉利翁氏菌屬(Gallionella)亞硝化單胞菌科(Nitrosomonadaceae)亞硝化葉菌屬(Nitrosolobus)亞硝化單胞菌屬(Nitrosomonas)亞硝化螺旋菌屬(Nitrosospira)螺菌科(Spirillaceae)螺菌屬(Spirillum)普羅卡桿菌目(Procabacteriales)*普羅卡桿菌科(Procabacteriaceae)*普羅卡桿菌屬(Procabacter)*紅環(huán)菌目(Rhodocyclales)紅環(huán)菌科(Rhodocyclaceae)固氮弓菌屬(Azoarcus)(Azonexus)(Azospira)(Azovibrio)(Dechloromonas)(Dechlorosoma)(Denitratisoma)(Ferribacterium)(Methyloversatilis)丙酸桿菌屬(Propionibacter)(Propionivibrio)(Quatrionicoccus)紅環(huán)菌屬(Rhodocyclus)(Shinella)(Sterolibacterium)陶厄氏菌屬(Thauera)(Uliginosibacterium)動膠菌屬(Zoogloea)21.3

δ-變形菌綱(Deltaproteobacteria)蛭弧菌目(Bdellovibrionales)(Bacteriovora